To state it a different way: yes, of course and without doubt their very presence affected biodiversity.

But you were talking about their practices, which tended towards custodial over exploitative. And overall these practices clearly supported biodiversity as a whole, otherwise we wouldn’t note the biodiversity of this region as anything special (see again the quote I took from your first article).

I apologize anyway for my slightly combative tone. I appreciate the resources you shared even if I haven’t had time to absorb them in full yet.