Checking the Firefox bugs on this, it seems they decided to replace the C++ libjxl with a rust version which is a WIP, to address security concerns with the implementation. All this started a few months ago.

Maybe the zen fork is a bit older and still using the C++ one?

... update. after reading the comments in the rust migration security bug, I saw they mentioned "only building in nightly for now"

I grabbed the nightly firefox, flipped the jxl switch, and it does indeed render fine, so I guess the rust implementation is functioning, just not enabled in stable.

... also, I see no evidence that it was ever enabled in the stable builds, even for the C++ version, so I'm guessing Zen just turned it on. Which... is fine, but maybe not very cautious.