Not all epigenetics is regulation of gene expression. The article says "these molecules transmit traits to offspring and that they can regulate embryonic development after fertilization" -- that's from the reporter, but I don't have faith that "transmit traits" is at all accurate--it certainly isn't true in the way that genes express traits. And then they quote an actual epigeneticist saying “We just don’t have really any understanding of how RNAs can do this, and that’s the hand-wavy part”