It would be interesting to see if left handers and right handers differ on this. I can adapt to any scheme with basically zero mental effort, and I've heard this is common in lefties (as I am). Stuff like "hold this up to a mirror for answer" never worked for me because I could read a whole page like that without noticing it's backwards. Da Vinci certainly had it with his inverted notebooks

Yeah, I always assumed my inversion preference was something to do with my left-handedness.

I'm guessing it should have been tested for in the study cited. Massive omitted variable bias if not.