So the queen can lay 3 types: hybrid female, Ibiricus male, structor male. Did they do karyotyping? Is it actually that the queen somehow removed its own genetic material from the nuclei or does it somehow get silenced when the M. structor genetic material is present in the nuclei (which is interesting by itself). Perhaps some kind of complex imprinting is happening.

Real life Zerg Queen..

Four types! You missed Ibiricus female, ie the next generation of queen